Anatoliy A. Khapugin

Managing Editor of the journal “Nature Conservation Research”

Academic degree, title, position: PhD (in Biology). Senior Researcher of the Joint Directorate of the Mordovia State Nature Reserve and National Park “Smolny", Researcher of the Tyumen State University.

Awards, membership in societies and editorial boards of journals: Member of the Russian Botanical Society. Anatoliy A. Khapugin consists in editorial boards of the journals Biodiversity Data Journal (Subject Editor), Natural Areas Journal (Subject Editor), Nature Conservation (Subject Editor), Check List (Subject Editor), Botanica Serbica (Editorial Board member), Biodiversity (Editorial Board member).

Research interests: plant ecology, botany, forest ecosystems, biological invasions, plant conservation.

Scientometric indicators: Web of Science (h=11, Documents by author: 53, Total citations: 372); Scopus (h=14, Documents by author: 70, Total citations: 590); Russian Index of Science Citation (RISC) (h=20, Documents by author: 224, Total citations: 2106).

Publication activity: Anatoliy A. Khapugin is author of more than 200 publications, including more than 130 articles in peer reviewed journals; he is an author (or co-author) of 18 monographs, 1 teaching aid.

Web-pages: ResearchGate, ORCID, Publons, Scopus, Russian Science Citation Index (RSCI).

Selected publications

  • Khapugin A.A., Senchugova M.A. 2022. The age structure of a Cypripedium guttatum (Orchidaceae) population in the Tyumen Region, Western Siberia // Journal of Wildlife and Biodiversity. Vol. 7(2). P. 35–44. DOI: 10.5281/zenodo.7178289
  • Khapugin A.A., Chugunov G.G. 2023. Population Status of a Regionally Endangered Plant, Lunaria rediviva (Brassicaceae), near the Eastern Border of Its Range // Biology. Vol. 12(6). Article: 761. DOI: 10.3390/biology12060761
  • Khapugin A.A., Esina I.G., Silaeva T.B. 2023. The Updated Check-List of Alien Plant Species in the Republic of Mordovia: A Path from 2010 to 2023 // Russian Journal of Biological Invasions. Vol. 14(3). P. 446–456.
  • Khapugin A.A., Kuzmin I.V., Ivanova L.A. 2021. Distribution of four alien plants in Tyumen Region (Western Siberia): contribution of citizen science and expert data. Wulfenia 28: 151–160.
  • Khapugin A.A., Kuzmin I.V. 2022. Data for Distribution of Vascular Plants (Tracheophytes) of Urban Forests and Floodplains in Tyumen City (Western Siberia). Data 7(12): 180. DOI: 10.3390/data7120180
  • Esina I.G., Khapugin A.A. 2022. To What Extent Are Protected Areas Freer of Alien Plants Than Managed Areas within Biodiversity Coldspots? A Case Study of the Mordovia State Nature Reserve, European Russia. Biology and Life Sciences Forum 15: 23. DOI: 10.3390/IECD2022-12416
  • Khapugin A.A., Kuzmin I.V., Ivanov L.A., Ronzhina D.A., Ivanova L.A. 2022. Environmental Preferences of an Invasive Plant Species, Bidens frondosa (Asteraceae), in European Russia and Western Siberia. Diversity 14(8): 598. DOI: 10.3390/d14080598
  • Ogurtsov S.S., Khapugin A.A., Zheltukhin A.S., Fedoseeva E.B., Antropov A.V., Delgado M.M., Penteriani V. 2022. Brown Bear Food-Probability Models in West-European Russia: On the Way to the Real Resource Selection Function. Forests 13(8): 1247. DOI: 10.3390/f13081247
  • Fardeeva M.B., Chizhikova N.A., Zheleznaya E.L., Khapugin A.A., Puchnina L.V., Suleimanova V.N., Ishmuratova M.M., Teteryuk L.V., Ilyina V.N., Urbanavichute S.P., Prokhorov V.E., Egorova N.Yu., Nabiullin M.I., Barlybaeva M.Sh., Kildiyarova G.N., Suyundukov I.V., Marakaev O.A. 2022. Demographic Structure of Cypripedium calceolus L. Populations in the European Part of Russia under Climate Change. Contemporary Problems of Ecology 15(4): 353–372. DOI: 10.1134/S1995425522040047
  • Sofronova E.V., Abdurakhmanova Z.I., Afonina O.M., Borovichev E.A., Boychuk M.A., Cherednichenko O.V., Czernyadjeva I.V., Doroshina G.Ya., Dulin M.V., Esina I.G., Fedosov V.E., Gadzhiataev M.G., Grishutkin O.G., Ignatov M.S., Ignatova E.A., Kessel D.S., Khapugin A.A., Kozhin M.N., Kuzmina E.Yu., Liksakova N.S., Maksimov A.I., Moshkovskii S.A., Popova N.N., Semenova N.A., Shchukina K.V., Sinichkina A.D., Suslova E.G., Tubanova D.Ya., Zheleznova G.V. 2021. New bryophyte records. 17. Arctoa 30: 458–470. DOI: 10.15298/arctoa.30.31
  • Seregin A.P., Bochkov D.A., Shner J.V., Garin E.V., Pospelov I.N., Prokhorov V.E., Golyakov P.V., Mayorov S.R., Svirin S.A., Khimin A.N., Gorbunova M.S., Kashirina E.S., Kuryakova O.P., Bolshakov B.V., Ebel A.L., Khapugin A.A., Mallaliev M.M., Mirvoda S.V., Lednev S.A., Nesterkova D.V., Zelenova N.P., Nesterova S.A., Zelenkova V.N., Vinogradov G.M., Biryukova O.V., Verkhozina A.V., Zyrianov A.P., Gerasimov S.V., Murtazaliev R.A., Basov Y.M., Marchenkova K.Yu., Vladimirov D.R., Safina D.B., Dudov S.V., Degtyarev N.I., Tretyakova D.V., Chimitov D.G., Sklyar E.A., Kandaurova A.N., Bogdanovich S.A., Dubynin A.V., Chernyagina O.A., Lebedev A.V., Knyazev M.S., Mitjushina I.Yu., Filippova N.V., Dudova K.V., Kuzmin I.V., Svetasheva T.Yu., Zakharov V.P., Travkin V.P., Magazov Y.O., Teploukhov V.Yu., Efremov A.N., Deineko O.V., Stepanov V.V., Popov E.S., Kuzmenckin D.V., Strus T.L., Zarubo T.V., Romanov K.V., Ebel A.L., Tishin D.V., Arkhipov V.Yu., Korotkov V.N., Kutueva S.B., Gostev V.V., Krivosheev M.M., Gamova N.S., Belova V.A., Kosterin O.E., Prokopenko S.V., Sultanov R.R., Kobuzeva I.A., Dorofeev N.V., Yakovlev A.A., Danilevsky Y.V., Zolotukhina I.B., Yumagulov D.A., Glazunov V.A., Bakutov V.A., Danilin A.V., Pavlov I.V., Pushay E.S., Tikhonova E.V., Samodurov K.V., Epikhin D.V., Silaeva T.B., Pyak A.I., Fedorova Y.A., Samarin E.S., Shilov D.S., Borodulina V.P., Kropocheva E.V., Kosenkov G.L., Bury U.V., Mitroshenkova A.E., Karpenko T.A., Osmanov R.M., Kozlova M.V., Gavrilova T.M., Senator S.A., Khomutovskiy M.I., Borovichev E.A., Filippov I.V., Ponomarenko S.V., Shumikhina E.A., Lyskov D.F., Belyakov E.A., Kozhin M.N., Poryadin L.S., Leostrin A.V. 2020. «Flora of Russia» on iNaturalist: a dataset. Biodiversity Data Journal 8: e59249. DOI: 10.3897/BDJ.8.e59249
  • Gammi L., Borah D., Taram M., Tag H., Khapugin A.A. 2020. Beccarinda tonkinensis (Gesneriaceae), a new record for India and its IUCN Red List status in India. Wulfenia 27: 139–144.
  • Khapugin A.A., Gafurova M.M., Suleymanova G.F., Bogdanov G.A., Chap T.F., Zenkova N.A., Belkovskaya T.P. 2020. The role of Russian federal protected areas in conservation of plant species included in the Red Data Book of Russian Federation (Volga Federal District). Wulfenia 27: 114–126.
  • Borah D., Das B., Tangjang S., Das A.P., Khapugin, A.A. 2021. Assessment of woody species diversity and composition along a disturbance gradient in Behali Reserve Forest of Biswanath district, Assam, India. Ecological Questions 32(1): 1–25. DOI: 10.12775/EQ.2021.009
  • Esina I.G., Khapugin A.A., Esin M.N., Sinichkina A.D., Silaeva T.B. 2021. Additions to the flora of the Republic of Mordovia, Russia. Contribuţii Botanice 56: 59–64. DOI: 10.24193/Contrib.Bot.56.6
  • Khapugin A. 2021. Environment Status Estimation of the Forest Communities Based on Floristic Surveys in the Mordovia State Nature Reserve, Russia. Forests 12(11): 1475. DOI: 10.3390/f12111475
  • Khapugin A.A., Kuzmin I.V., Ivanova L.A. 2021. Distribution of four alien plants in Tyumen Region (Western Siberia): contribution of citizen science and expert data. Wulfenia 28: 151–160.
  • Khapugin A.A., Sołtys-Lelek A., Fedoronchuk N.M., Muldashev A.A., Agafonov V.A., Kazmina E.S., Vasjukov V.M., Baranova O.G., Buzunova I.O., Teteryuk L.V., Dubovik D.V., Gudžinskas Z., Kukk T., Kravchenko A.V., Yena A.V., Kozhin M.N., Sennikov A.N. 2021. Taxon-level assessment of the data collection quality in Atlas Florae Europaeae: insights from the case of Rosa (Rosaceae) in Eastern Europe. Nordic Journal of Botany 39(9): NJB12566. DOI: 10.1111/njb.03289
  • Łobas Z., Khapugin A., Żołubak E., Jakubska-Busse A. 2021. The Epipactis helleborine Group (Orchidaceae): An Overview of Recent Taxonomic Changes, with an Updated List of Currently Accepted Taxa. Plants 10(9): 1839. DOI: 10.3390/plants10091839
  • Ronzhina D.A., Ivanova L.A., Ivanov L.A., Khapugin A.A. 2021. Bidens frondosa (Asteraceae), a new alien invasive plant species in the Kurgan Region (Russia). Contribuţii Botanice 56: 53–58. DOI: 10.24193/Contrib.Bot.56.5
  • Ruchin A.B., Alekseev S., Khapugin A., Esin M. 2021. Fauna and Species Diversity of Ground Beetles (Coleoptera, Carabidae) in Meadows. Entomology and Applied Science Letters 8(3): 28–39. DOI: 10.51847/NV94GSLSKN
  • Ruchin A.B., Egorov L.V., Khapugin A.A. 2021. Seasonal activity of Coleoptera attracted by fermental crown traps in forest ecosystems of Central Russia. Ecological Questions 32(1): 1–21. DOI: 10.12775/EQ.2021.004
  • Ruchin A.B., Egorov L.V., Khapugin A.A. 2021. Usage of Fermental Traps for the Study of the Species Diversity of Coleoptera. Insects 12(5): 407. DOI: 10.3390/insects12050407
  • Ruchin A.B., Egorov L.V., MacGowan I., Makarkin V.N., Antropov A.V., Gornostaev N.G., Khapugin A.A., Dvořák L., Esin M.N. 2021. Post‑fire insect fauna explored by crown fermental traps in forests of the European Russia. Scientific Reports 11: 21334. DOI: 10.1038/s41598-021-00816-3
  • Sofronova E.V., Abdurakhmanova Z.I., Afonina O.M., Borovichev E.A., Boychuk M.A., Cherednichenko O.V., Czernyadjeva I.V., Doroshina G.Ya., Dulin M.V., Esina I.G., Fedosov V.E., Gadzhiataev M.G., Grishutkin O.G., Ignatov M.S., Ignatova E.A., Kessel D.S., Khapugin A.A., Kozhin M.N., Kuzmina E.Yu., Liksakova N.S., Maksimov A.I., Moshkovskii S.A., Popova N.N., Semenova N.A., Shchukina K.V., Sinichkina A.D., Suslova E.G., Tubanova D.Ya., Zheleznova G.V. 2021. New bryophyte records. 17. Arctoa 30: 458–470. DOI: 10.15298/arctoa.30.31
  • Verkhozina A.V., Biryukov R.Yu., Bogdanova E.S., Bondareva V.V., Chernykh D.V., Dorofeev N.V., Dorofeyev V.I., Ebel A.L., Efimov P.G., Efremov A.N., Erst A.S., Fateryga A.V., Gamova N.S., Glazunov V.A., Gudkova P.D., Juramurodov I.J., Kapitonova O.А., Kechaykin A.A., Khapugin A.A., Kosachev P.A., Krupkina L.I., Kulagina M.A., Kuzmin I.V., Lian L., Koychubekova G.A., Lazkov G.A., Luferov A.N., Mochalova O.A., Murtazaliev R.A., Nesterov V.N., Nikolaenko S.A., Novikova L.A., Ovchinnikova S.V., Plikina N.V., Saksonov S.V., Senator S.A., Silaeva T.B., Suleymanova G.F., Sun H., Tarasov D.V., Tojibaev K.Sh., Vasjukov V.M., Wang W., Zibzeev E.G., Zolotov D.V., Zykova E.Yu., Krivenko D.A. 2021. Findings to the flora of Russia and adjacent countries: New national and regional vascular plant records, 3. Botanica Pacifica 10(1): 85–108. DOI: 10.17581/bp.2021.10110
  • Ruchin A.B., Carr J.F., Dvořák L. Esin M.N., Khapugin A.A. 2020. Pseudotephritis millepunctata (Hennig, 1939) (Diptera Ulidiidae) – new species in European fauna. Redia 103: 25–27. DOI: 10.19263/REDIA-103.20.05
  • Khapugin A.A. 2020. A global systematic review on orchid data in Protected Areas. Nature Conservation Research 5(Suppl.1): 19–33. DOI: 10.24189/ncr.2020.019
  • Khapugin A.A., Silaeva T.B. 2020. The arrangement of threatened plants in Mordovia: the role of biodiversity research centers. Écoscience 27(3): 157–164. DOI: 10.1080/11956860.2020.1753293
  • Khapugin A.A., Kuzmin I.V., Silaeva T.B. 2020. Anthropogenic drivers leading to regional extinction of threatened plants: insights from regional Red Data Books of Russia. Biodiversity and Conservation 29(8): 2765–2777. DOI: 10.1007/s10531-020-02000-x
  • Verkhozina A.V., Chernysheva O.A., Ebel A.L., Erst A.S., Dorofeev N.V., Dorofeyev V.I., Grebenjuk A.V., Grigorjevskaja A.Y., Guseinova Z.A., Ivanova A.V., Khapugin A.A., Korolyuk A.Y., Korznikov K.A., Kuzmin I.V., Mallaliev M.M., Murashko V.V., Murtazaliev R.A., Popova K.B., Safronova I.N., Saksonov S.V., Sarajeva L.I., Senator S.A., Troshkina V.I., Vasjukov V.M., Wang W., Xiang K., Zibzeev E.G., Zolotov D.V., Zykova E.Y., Krivenko D.A. 2020. Findings to the flora of russia and adjacent countries: New national and regional vascular plant records, 2. Botanica Pacifica 9(1): 139–154. DOI: 10.17581/BP.2020.09115
  • Linnik E.V., Khapugin A.A. 2020. Status of Tilia maximowicziana (Malvaceae s.l.) in Russia. Nature Conservation Research 5(2): 97–102. DOI: 10.24189/ncr.2020.007
  • Chugunov G.G., Khapugin A.A. 2020. Population status of Bupleurum aureum (Apiaceae), a Critically Endangered plant species in a region of European Russia. Ecological Questions 31(1): 45–56. DOI: 10.12775/EQ.2020.006
  • Ruchin A.B., Egorov L.V., Khapugin A.A., Vikhrev N.E., Esin M.N. 2020. The use of simple crown traps for the insects collection. Nature Conservation Research 5(1): 87–108. DOI: 10.24189/ncr.2020.008
  • Ruchin A.B., Khapugin A.A. 2019. Red Data Book Invertebrates in a Protected Area of European Russia. Acta Zoologica Academiae Scientiarum Hungaricae 65(4): 349–370. DOI: 10.17109/AZH.65.4.349.2019
  • Khapugin A.A. 2019. Benefits from visualization of environmental factor gradients: a case study in a protected area in central Russia. Revista Chapingo Serie Ciencias Forestales y del Ambiente 25(3): 383–398. DOI: 10.5154/r.rchscfa.2019.01.004
  • Le Roux J.J., Hui C., Castillo M.L., Iriondo J.M., Keet J.H., Khapugin A.A., Médail F., Rejmánek M., Theron G., Yannelli F.A., Hirsch H. 2019. Recent Anthropogenic Plant Extinctions Differ in Biodiversity Hotspots and Coldspots. Current Biology 29(17), pp. 2912-2918.e2. DOI: 10.1016/j.cub.2019.07.063
  • Khapugin A.A. 2019. A global systematic review of publications concerning the invasion biology of four tree species. Hacquetia 18(2): 233–270. DOI: 10.2478/hacq-2019-0005
  • Ruchin A.B., Alekseev S.K., Khapugin A.A. 2019. Post-fire fauna of carabid beetles (Coleoptera, Carabidae) in forests of the Mordovia State Nature Reserve (Russia). Nature Conservation Research 4(Suppl.1): 11–20. DOI: 10.24189/ncr.2019.009
  • Ruchin A.B., Antropov A.V., Khapugin A.A. 2019. Distribution, abundance, and habitats of rare species Parnopes grandior (Pallas 1771) (Hymenoptera, Chrysididae) in Mordovia and adjacent regions, Russia. Biodiversitas 20(2): 303-310. DOI: 10.13057/biodiv/d200201
  • Khapugin A.A. Kunaeva E.N., Silaeva T.B. 2018. Extreme drought conditions can drastically change age structure of a plant population: case study of Jacobaea paludosa subsp. lanata (Family: Asteraceae) in the Republic of Mordovia (Russia). Journal of Bombay Natural History Society 115: 21–24. DOI: 10.17087/jbnhs/2018/v115/116395
  • Khapugin A., Senchugova M. 2018. The floristic lists as a source to characterize environment conditions of habitats using phytoindication methods: A case study for Iris aphylla (Iridaceae) and Lilium martagon (Liliaceae) in central Russia. Arnaldoa 25(1): 75–86. DOI: 10.22497/arnaldoa.251.25104
  • Khapugin A.A. 2018. Population characteristics of Silaum silaus (L.) Schinz & Thell. (Apiaceae) in Mordovia, a highly threatened plant species at the northern limit of its range. Journal of Threatened Taxa 10(1): 11147–11155. DOI: 10.11609/jott.3476.10.1.11147-11155
  • Khapugin A.A., Chugunov G.G., Vargot E.V. 2017. Cypripedium calceolus (Orchidaceae) in Central Russia: a case study for its populations in two Protected Areas in the Republic of Mordovia (Russia). Lankesteriana 17(3): P. 403–417. DOI: 10.15517/lank.v17i3.31577
  • Khapugin A.A., Korochkina A.M. 2017. Population-based study of Platanthera bifolia in the Mordovia State Nature Reserve (Republic of Mordovia, Central Russia). Orchidee Spontanee d'Europa – European Native Orchids 60(2): 276–294.
  • Khapugin A.A. 2017. A record of Silene viscaria (L.) Jess. (Caryophyllaceae) with achromatic flowers in the Mordovia State Nature Reserve (Central Russia). Annales Universitatis Paedagogicae Cracoviensis Studia Naturae 2. DOI: 10.24917/25438832.2.8
  • Khapugin A.A. 2017. Hieracium sylvularum (Asteraceae) in the Mordovia State Nature Reserve: invasive plant or historical heritage of the flora? Nature Conservation Research 2(4): 40–52. DOI: 10.24189/ncr.2017.013
  • Khapugin A.A., Silaeva T.B., Vargot E.V., Chugunov G.G., Grishutkina G.A., Grishutkin O.G., Pismarkina E.V., Orlova Ju.S. 2017. Estimation of taxa included in the first volume of the Red Data Book of the Republic of Mordovia (Russia) using the IUCN Red List Categories and Criteria. Nature Conservation Research. Vol. 2(Suppl. 1). P. 164–189. DOI: 10.24189/ncr.2017.004 [In Russian]
  • Khapugin A.A., Silaeva T.B., Vargot E.V., Chugunov G.G. 2017. IUCN guidelines using for assessment of plants from the Red Book of Russian Federation at regional level: a case study for the Republic of Mordovia (Russia). Hacquetia 16 (1): 19–33. DOI: 10.1515/hacq-2016-0012.
  • Khapugin A.A., Chugunov G.G., Vargot E.V., Silaeva T.B. 2017. Vascular plants at the protected areas network of the Republic of Mordovia: present status and prospects. In: S.A. Mukul, A.Z.M.M. Rashid (Eds.): Protected Areas: Policies, Management and Future Directions. USA: Nova Science Publishers, Inc. P. 203–231.
  • Khapugin A.A., Kunaeva E.N. 2017 Abundance dynamics of endangered plant species Galatella rossica (Asteraceae) in the Republic of Mordovia (Central Russia): short report. Arnaldoa 24(1): 239–246. DOI: 10.22497/arnaldoa.241.24109
  • Khapugin A.A., Silaeva T.B., Utorova Yu.N. 2016. Three Maples (Acer L., Aceraceae Juss.) in the Republic of Mordovia, Russian Federation. Annales Universitatis Paedagogicae Cracoviensis Studia Biologica 1: 129–143.
  • Khapugin A.A., Chugunov G.G., Silaeva T.B., Kunaeva E.N. 2016. Neottianthe cucullata (L.) Schltr. (Orchidaceae Juss.), an endangered orchid in Central Russia. Wulfenia 23: 189–202.
  • Khapugin A.A., Vargot E.V., Chugunov G.G., Shugaev N.I. 2016. Invasion of alien plants in fire-damaged forests at Southern boundary of the taiga zone. Forest Systems 25(3): eSC13. DOI: 10.5424/fs/2016253-09461.
  • Khapugin A.A., Silaeva T.B., Semchuk A.A., Kunaeva E.N. 2016. Populations of Orchis militaris L., Epipactis palustris (L.) Crantz and Malaxis monophyllos (L.) Sw. in the Republic of Mordovia (Central Russia). Biodiversity: Research and Conservation 42: 33–40. DOI: 10.1515/biorc-2016-0012
  • Khapugin A.A., Vargot E.V., Chugunov G.G. 2016. Vegetation recovery in fire-damaged forests: a case study at the southern boundary of the taiga zone. Forestry Studies 64: 39–50. DOI: 10.1515/fsmu-2016-0003
  • Gladunova N.V., Khapugin A.A., Vargot E.V. 2016. Bidens frondosa L. (Asteraceae) in the Republic of Mordovia (Russia). Russian Journal of Biological Invasions 7(2): 129–136. DOI: 10.1134/S2075111716020053.
  • Khapugin A.A., Chugunov G.G. 2015. Two populations of Lunaria rediviva L. (Cruciferae Juss.) at the eastern edge of its range. Wulfenia 22: 83–94.
  • Khapugin A.A., Semchuk A.A., Silaeva T.B., Chugunov G.G. 2014. A comparative characteristics of Cypripedium calceolus L. (Orchidaceae, Monocotyledones) coenopopulations in the Republic of Mordovia. Povolzhskiy Journal of Ecology 3: 403–410. [In Russian]
  • Gladunova N.V., Vargot E.V., Khapugin A.A. 2014. Oenothera biennis L. (Onagraceae) in the Republic of Mordovia (Russia). Russian Journal of Biological Invasions 5(1): 12–17. DOI: 10.1134/S2075111714010044.
  • Utorova Yu.N., Khapugin A.A., Silaeva T.B. 2014. About Ecology of Acer campestre L. (Aceraceae) on North-Eastern Limit of the Range. Environment and Ecology Research 2(1): 8–13. DOI: 10.13189/eer.2014.020102
  • Khapugin A.A., Vargot E.V., Chugunov G.G., Dementeva A.E. 2013. Additions and Notes to the Alien Flora of the Mordovia State Nature Reserve. Russian Journal of Biological Invasions 4(3): 200–207. DOI: 10.1134/S2075111713030041.
  • Khapugin A.A., Buzunova I.O. 2013. The synopsis of section Caninae DC. of genus Rosa L. (Rosaceae) in the flora of the Moksha river basin. Novosti sistematiki vysšich rastenij 44: 135–145. [In Russian]
  • Khapugin A.A., Silaeva T.B. 2013. The Cover of Steppe Vascular Plants by the Protected Area Network of the Republic of Mordovia (Russia). In: M.B. Morales Prieto, J. Traba Diaz (Eds.): Steppe Ecosystems: Biological Diversity, Management and Restoration. New York, Nova Science Publishers, Inc. P. 211–220.
  • Khapugin A.A. 2012. About finding of Rosa glauca Pourr. (Rosaceae) in the Republic of Mordovia. Russian Journal of Biological Invasions 3(1): 56–57. DOI: 10.1134/S2075111712010043.
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